Adventitious roots (Dennis Murengi)
Anticlinal wall - Tyler Parnall
Apical meristem
Casparian strip- Liz Foster
Cortex
Endodermis
Epidermis- Jessi.E.W
Exodermis - Jackie Taylor
Feeder roots - Amanda Miller
Fibrous root system - Kevin Christie
Frankia
Germination - Krista Neufeld
Ground meristems- Nicole L.
Hypocotyl
Imbibition - Tori Waites
Lateral meristem
Meristematic region-Danielle Dorink
Mucigel
Mycorrhizae-Danielle Dornik
Nitrogen fixation - Russ Fountain
Periclinal wall
Pericycle
Primary meristems
Primary phloem (Dennis Murengi)
Primary xylem - Tyler Parnall
Procambium
Promeristem - Morgan Traverse
Protoderm - Kirsten Jorgensen
Protophloem - Kristie S
Protoxylem - Stephanie Lauer
Radial wall
Radicle - Amanda Miller
Rays
Root hairs
Root nodules - Kevin Christie
Stele
Suberin
Tap root system
Transverse wall
Turgor pressure - Tori Waites
Vascular cambium
Definitions (in alphabetical order) Adventitious roots (Dennis Murengi) - A root that grows from somewhere other than the primary root, for example, from stems or leaves. (www.en.mimi.hu)
http://en.mimi.hu/gardening/adventitious_roots.html Most commonly, adventitious roots arise out of stems, originating via cell divisions of the stem cortex or less often from auxiliary buds hidden in the bark. In some plants plant leaves are encouraged to form adventitious roots. Examples include: adventitious roots of a palm and of an aquatic plant that has unattached roots in moving water. (www.botgard.ucla.edu)
http://www.botgard.ucla.edu/html/botanytextbooks/generalbotany/typesofroots/index.html
The image above shows adventitious roots growing from the stem of a plant. (www.lexic.us)
http://www.lexic.us/definition-of/adventitious_rootCasparian Strip: The casparian strip is a band of primary cell walls that contain suberin (which is a fatty substance) on the anticline walls of the endodermal cells that stop water and minerals from seeping in between the cells and therefore forces the water and minerals to enter the endodermal cells and therefore be filtered before it enters the vascular cylinder. The main component in the casparian strip, suberin is named after the cork oak, as suberin is also found in cork. The casparian strip is named after the 19th century German scientist that discovered it, Robert Caspary.
http://faculty.clintoncc.suny.edu/faculty/michael.gregory/files/Bio%20102/Bio%20102%20lectures/Plant%20Structure/plant%20structure.htm (Clinton Community College),
http://www.daviddarling.info/encyclopedia/C/casparian_strip.html (David Darling Encyclopedia),
http://www.biochem.northwestern.edu/holmgren/Glossary/Definitions/Def-C/Casparian_strip.html (Northwestern)
Liz Foster
Enhancement- A band like region of primary wall containing suberin and lignin. It is found in anticlinal-radial and transverse-wall of the endodermal and the exodermal cells. Only angiosperms have casparian strips in the exodermis.-Danielle Dornik
(Biology of Plants, 7th edition, Peter H. Raven G-4)
Endodermis – endo meaning inner dermis meaning skin The endodermis is a thin layer of cells bounding the stele and separating it from the cortex. The cells, surrounds the central core of the vascular tissue, and which helps to regulate the flow of water and dissolved substances. The endodermis is one of the most important adaptations of terrestrial plants. (Encyclopaedia Britannica and Merriam-Webster)

www.apsnet.org www.botany.hawaii.edu/Enhancement:
Cells in the endodermis are waterproof and fit together very tightly with no intercellular spaces. These features allow the endodermis to be a highly effective barrier between vascular tissue and other cells. (Plant anatomy laboratory: http://www.sbs.utexas.edu/mauseth/weblab/webchap3par/3.4-2.htm). Katherine Loewen
Exodermis: (Jackie Taylor) The outer layer, one or more cells in depth, of the cortex in some roots; these cells are characterized by Casparian strips within the radial and transverse walls. The exodermis is located just below the epidermis.It is developed from the outermost layer or layers of cells of the cortex. The majority of angiosperm roots also have an exodermis with a Casparian stripes. Following development of Casparian strips, a suberin lamella is deposited on all walls of the exodermis. The exodermis reduces water loss for the root to the soil and provides a defense against attack by microorganisms. Under stress the exodermis develops closer to the root tip.
Enhancement (Exodermis) Russ Fountain
Below is a picture of a
Smilax (greenbrier) root. Note the distinctive endodermis. The outermost layer of the cortex is differentiated as an
exodermis, similar in appearance and function to the endodermis.

(“Roots play a vital role in plants, which is to take up water and nutrients from the soil.” -
http://www.northern.edu/ramsayj/etechWeb/roots.htm)
(Springer Link - http://www.springerlink.com/content/3rehrkkwy3k1g1b1/) (http://trc.ucdavis.edu/biosci10v/bis10v/week8/casparian.gif) (Biology of Plants, 7th edition, Peter H. Raven, Glossary pg. G-9)Feeder roots: Fine roots that activly take up water and minerals through root hairs, they are generally found within the top meter of soil. (The American Heritage Science Dictionary By Houghton Mifflin Company http://books.google.com/books?id=yKUagx8PB_EC&pg=PA544&lpg=PA544&dq=feeder+roots+dictionary&source=web&ots=KYqoT9BVK0&sig=mXMISi51qOMxEKcQhXwUFdJYsp8#PPA544,M1 Feeder roots have a symbiotic relationship with mycorrhizal fungi, the fungi are attached to these small feeder roots. (Encyclopedia Britannica Online http://www.britannica.com/eb/article-9054526/mycorrhiza)Amanda MEnhacement: Liz Foster
Note the small white feeder roots. http://pubs.caes.uga.edu/.../B813-w.htg/B813-7b.jpg, (University of Georgia) Feeder Roots (enhancment): Feeder roots are responsible for the majority of absorbtion of nutrient of a tree. even though feeder roots are only 1/6 inches in diameter, they make up a large portion of a trees root system surface area. These smaller roots grow outward and predominantly upward from the large roots near the soil surface, where minerals, water and oxygen are relatively abundant. Under normal conditions the feeder roots die and are replaced on a regular basis. (
http://www.ext.colostate.edu/pubs/garden/02926.html) - VanessaV
Fibrous Root System: Root system in plants which no one root is more prominent than the others. Typically shallower than tap root system and the tenacity with which they cling to soil particle make such plants especially well suited as ground cover for the prevention of soil erosion. (
Biology of Plants, 7th edition, Peter H. Raven, pg. 528 – 529)

Fibrous root system of a
hydroponically grown plant.
http://commons.wikimedia.org/wiki/Image:Roots.JPGKevin Christie
Enhancement:
A
fibrous root system (sometimes also called
adventitious root system) is the opposite of a taproot system. It is usually formed by thin, moderately branching roots growing from the stem. A fibrous root system is universal in monocotyledonous plants and ferns, and is also common in dicotyledonous plants. Most trees begin life with a taproot, but after one to a few years change to a wide-spreading fibrous root system with mainly horizontal surface roots and only a few vertical, deep anchoring roots. Weeds have fibrous root systems, and that is why some are so hard to pull from the ground. Also, if extra roots remain in the soil, the weeds may grow back again.
http://en.wikipedia.org/wiki/Fibrous_root_systemKrista Neufeld
Enhancement: Grasses and other monocotyledons have a fibrous root system, characterized by a mass of roots of about equal diameter. This network of roots does not arise as branches of the primary root but consists of many branching roots that emerge from the base of the stem.(Britannica On-line)Amanda LastGermination - This is the process whereby growth emerges from a period of dormancy. The most common example of germination is the sprouting of a seedling from a seed of an angiosperm or gymnosperm. Germination is the first stage of the making of the seedling. The seed of a higher plant is a small package produced in a flower or cone containing an embryo and stored food reserves. Under favorable conditions, the seed begins to germinate, and the embryonic tissues resume growth, developing towards a seedling. Below is an image of a sunflower seedling just 3 days after being planted. (http://en.wikipedia.org/wiki/Image:Sunflower_seedlings.jpg(http://en.wikipedia.org/wiki/Germination)
Krista Neufeld
Germination Enhancement: The three most important factors that germination is dependent on are water, oxygen, and temperature. Some seeds require exposure of light. Germination is not possible until the seed imbibes enough water, the surrounding soil contains enough oxygen for aerobic pathway, and the specific temperature range for the particular species is met. (Biology of Plants, Raven, p 504) Kirsten Jorgensen
Ground Meristems - Nicole L. The primary meristem, or meristematic tissue, that gives rise to the ground tissue. (biology of Plants, 7th edition, pg G-10) The primary meristem in vascular plants that gives rise to the nonvascular tissues, such as cortex, pericycle, and pith. Within the seeds of angiosperms, it surrounds the procambium. The primary meristem that differentiates into the pith and cortex (http://dictionary.reference.com/browse/Ground Meristem)
Enchanement: Nick N
Below is a diagram of the root apical meristem, the ground meristem is located around the outside of the procambium.
www.answers.com/topic/apical-meristemImbibition: Derived from the Latin word “imbibere” which means “to drink”, imbibition refers to the movement of water into a seed. This absorption of water into the colloidal material of cells in a seed causes swelling and increases Turgor pressure. Imbibition is critical in seed germination because without the subsequent swelling the embryo would not be able to escape the seed coat. The excessive presence of water also initiates the physiological changes that lead to germination.
(Biology of Plants, 7th Edition, Peter H. Raven, pg 76, G-12) (Koning, Ross E. "Home Page for Ross Koning". Plant Physiology Information Website. 1994. http://plantphys.info/seedg/seed3.html) Picture from: http://plantphys.info/seedg/seed3.html– Tori Waites
Meristematic Region-Danielle Dornik-(merizein- to divide)- Embryonic tissue regions, primarily concerned with formation of new cells. The meristematic region consists of meristematic tissue:
- Root Cap
- Promeristem
- Root apical meristem(protoderm, ground meristem, procambium)
- Primary meristems
The meristematic region is prtected by the root cap. The cells are very active and divide rapidly to form new cells which differentiate later to form more specialized root tissue. The cells of this region also replace the cells rubbed off from the root cap and provide additional cells for the next region.
(Biology of Plants, Peter H Raven G-14)
(
http://www.botany.uwc.ac.za/sci_ed/grade10/anatomy/roots.htm)
Enhancement (Stephanie Lauer): this is a cross section of a root and a shoot which shows the different meristematic tissues, where they are located and what makes them up.
http://cache.eb.com/eb/image?id=5599&rendTypeId=4 Mucigel: Mucigel is a slimy sheath covering the surface of roots (Raven G-14). It is part of the rootcap that penetrates through the soil. As most soil is compact and coarse, mucigel acts as a lubricant to aid the root in pushing through the soil. It accumulates in the golgi vesicles and is secreted through the root cap cells (Raven pg. 530). Due to its sticky nature, soil will bind and this benefits the root through water and nutrient absorption. It also encourages the growth of fungi and bacteria that are able to correct nitrogen in the soil. http://dictionary.reference.com/browse/mucigel --John
Enhancement (Nicole L)
Image from http://www.vitalearth.com/images/news/rizo-root.gif Image shows the mucigel around the outside surface of the root
Enhancement: Some functions of Mucigel are: Nutrient absorption-Carboxyl groups in mucigel influence ion uptake, and organic acids in mucigel make certain ions more available to plants. Also, fatty acids, lectins, and sterols in mucigel may help establish beneficial symbioses with soil microbes. Protection-It protects roots from desiccation and contains compounds that diffuse into the soil and inhibit growth in other roots. (Plant Physiology, www.cartage.org.lb/en/themes/sciences/botanicalsciences/PlantsStructure/Roots/Roots.htm) Mark Haywood-Farmer
Mycorrhizae- Myco-fungus rhiza-root -A symbiotic association between certain fungi and plant roots; characteristics of most vascular plants. (Biology Of Plants, Peter H. Raven,p262)
3 types:
Ectomycorrhizae-fungus forms sheath around root. Penetrates between cells of cortex to facilitate nutrient exchange
ectomycorrhizas of Suillus subluteus with Pinus resinosa (the branched ends of the short lateral roots)
Endomycorrhizae- vascular arbuscular mycorrhizae (VAM) Penetrates the corticle cells, doesn’t penetrate cell membrane of form sheath around root. More common than ectomycorrhizae.
colonization of a root by an endomycorrhizal fungus. Note hyphae, arbuscules and vesicles. (see Fig 21 in Brundrett et al. 1985 Can. J. Bot 63: 184)Orchid Mycorrhizae-special type. a symbiotic relationship between the roots of plants of the family Orchidaceae and a variety of fungi

r
oot cells of Goodyera oblongifolia containing coils (pelotons) of hyphae (probably of a mycorrhizal Rhizoctonia).Thomas J. Volk, University of Wisconsin-La Crosse (http://www.uwlax.edu/BIOLOGY/VOLK/FUNGI3/index.htm)
all pictures and descriptions from
http://www.mycolog.com/chapter17.htm-Danielle Dornik
Enhancement: (Nikki Heim) | About 95% of the world’s land plants form the mycorrhizal relationship in their native habitats. It is estimated that mycorrhizal fungal filaments can make up hundreds to thousands more soil volume compared to roots alone. Mycorrhizal fungus benefit plants by improving nutrient and water uptake for plant growth, root growth, resistance to disease, and reduced drought stress. |
| |
References: Mycorrhizal Application, Inc. Online. Access at http://www.mycorrhiza.com/index)
Mycorrhizae Enhancement 2: To emphasize that the relationship is mutualistic, the plant provides the fungi with energy. (The Ecology of Mycorrhizae, Allan, p.1) kirsten jorgensen
Nitrogen fixation (Russ Fountain)
The process by which nitrogen in the atmosphere is converted into nitrogenous compounds by the action of micro-organisms.(“nitrogen fixation” The Free Dictionary.
http://encyclopedia.farlex.com/nitrogen+fixation) Symbiotic nitrogen fixation occurs in plants that harbor nitrogen-fixing bacteria within their tissues. The best-studied example is the association between legumes and bacteria in the genus Rhizobium. Each of these is able to survive independently (soil nitrates must then be available to the legume), but life together is clearly beneficial to both. Only together can nitrogen fixation take place. (Symbiotic Nitrogen Fixation -
http://users.rcn.com/jkimball.ma.ultranet/BiologyPages/N/NitrogenFixation.html)
Enhancement (Sarah Slater): Nitrogen is one of the most abundant elements in the earth’s atmosphere and is an essential part of plant growth. When land is used for agriculture the soil is drained of nitrogen, thus farmers rotate crops with nitrogen fixing plants. Examples of plants commonly used for nitrogen fixation are legumes, which include alfalfa.
Alnus Rubra is an example of a tree that also fixes nitrogen. In the picture below you can see the root nodules, which is where the nitrogen fixation takes place.
Picture found at: Reed College <http://academic.reed.edu/biology/Nitrogen/Nfix1.html> Biology of Plants. Raven. G-14
American Chemical Society http://pubs.acs.org/subscribe/journals/esthag-w/2007/june/science/rr_nitrate.html
Periclinal Wall: Periclinal wall is literally translated as a wall that is parallel to the surface (Raven G-17). It is the exterior wall of epidermal cells. It is also known as anticlinal wall. http://jcs.biologists.org/cgi/reprint/48/1/193.pdf --John Pericycle: A thin layer of parenchymatous or sclerenchymatous cells that surrounds the stele in most vascular plants. (Merriam-Webster Online,
http://www.m-w.com/dictionary/pericycle).
It is plant tissue characteristic of roots: the outer layer of plant tissue surrounding the inner core of the roots and stems of plants stele that conducts moisture and nutrients around the plant(MSN Encarta Encyclopedia,
http://encarta.msn.com/encnet/refpages/search.aspx?q=Pericycle ). Lee Williams
Primary phloem (Dennis Murengi) – This is defined as the phloem of the primary plant body. (arabidopsis.info)
http://arabidopsis.info/cgibin/ontology/nasc_po/go.cgi?action=query&query=phloem&search_constraint=terms&view=query Primary phloem, are formed by the apical meristems of root and shoot tips; it may be either protophloem, the cells of which are matured before elongation (during growth) of the area in which it lies, or metaphloem, the cells of which mature after elongation. Sieve tubes of protophloem are unable to stretch with the elongating tissues and are torn and destroyed as the plant ages. (www.britannica.com)
http://www.britannica.com/eb/article-9059738/phloemProcambium- A primary meristematic tissue that gives rise to primary vascular tissues (Raven, Biology of Plants, Pg. 498). The part of the meristem of a plant that forms cambium and primary vascular tissue (Britannica Online Encyclopedia).
As the long band of red procambium continues to divide, some of their progeny cells will become vascular tissue-- xylem and phloem (Plant Anatomy Laboratory, James D. Mauseth,
University of Texas). Tessa JPromeristem: The initiating cells in the apical meristem (Biology of Plants, Peter H. Raven, pg.531) . These are the cells that are closest to the root tip and are responsible for producing the cells for the rest of the meristem (http://www.sbs.utexas.edu/mauseth/weblab/webchap6apmer/6.2-2.htm).
-Morgan TraverseEnhancement (Daniel Struthers): Promeristems are continually in the process of dividing cells and are present for the entire life of the root. When the root cap wears away due to root movement through the soil it is continuously being built back by the cells created from the promeristem. Source: The American Journal of Botany (online). http://www.jstor.org/view/00029122/di001736/00p0049a/0
Protoderm: Formed by periclinal divisions, the protoderm is the precursor of the epidermis. It is the primary meristematic tissue that develops into dermal tissue.(Biology of Plants, Raven, p 498), (Plant Anatomy, 4th Ed., Fahn, p 539)
http://www.northern.edu/ramsayj/etechWeb/images/roots/zea_rtls1B_copy.JPG. Retrieved November 26, 2007.text and photo-Kirsten jorgensen
Enhancement: (Liz Foster) The protoderm is the outermost layer of cells across the apical meristem and the leaf primordium.

The light mauvey purple is the protoderm.
www.sbs.utexas.edu/mauseth/weblab/webchap6apmer/6.2-2.htm (University of Texas)
Protophloem (Kristie Seaborn) the first-formed elements of the primary phloem, the cells of which mature before the elongation of the surrounding plant tissue; metaphloem (Oxford English Dictionary). The protophloem mother cell (PMC) undergoes two successive asymmetrical divisions: the first one gives rise to a smaller cell that will differentiate into a companion cell, and a larger one that divides again asymmetrically yielding another companion cell and a protophloem sieve element (http://jcs.biologists.org/cgi/content/abstract/75/1/165).
Enhancement: By Conyr Aird image 1

Image 2
These two images are photos of early vascular cell development and clearly depicts from left to right within the trace protoxylem, procambium, and protophloem in the first picture and are labeled in the second. ( Image 1 retrieved from The Stems webpage at the addresshttp://images.google.ca/imgres?imgurl=http://sols.unlv.edu/Schulte/Anatomy/Stems/PineVascClose.jpg&imgrefurl=http://sols.unlv.edu/Schulte/Anatomy/Stems/Stems.html&h=819&w=1024&sz=353&hl=en&start=9&um=1&tbnid=gzi0YovZ19BqUM:&tbnh=120&tbnw=150&prev=/images%3Fq%3Dprotophloem%26svnum%3D10%26um%3D1%26hl%3Den image 2 retrieved from botany Hawaii at the addresshttp://images.google.ca/imgres?imgurl=http://www.botany.hawaii.edu/faculty/webb/BOT410/Phloem/ProcambiumLSLab.jpg&imgrefurl=http://www.botany.hawaii.edu/faculty/webb/BOT410/Phloem/Phloem-1.htm&h=344&w=300&sz=34&hl=en&start=6&um=1&tbnid=LLe_nBJLvUw9LM:&tbnh=120&tbnw=105&prev=/images%3Fq%3Dprotophloem%26svnum%3D10%26um%3D1%26hl%3Den)
Protoxylem (Stephanie Lauer)– “proto” meaning first but not fully functional. The part of the primary xylem that consists of tracheids (Ross E. Koning 1994) which develops first and matures during elongation of roots, stems and leaves of the plant that it is found in (Raven, Biology of Plants, pg 517) . Koning, Ross E. 1994. "Root Vocabulary". Plant Physiology Website. http://plantphys.info/Plant_Biology/rootvocab.html (November 28, 2007)
Enhancement: (Dennis Murengi)-The image below shows a cross section of a root’s region of elongation. The location of the protoxylem is clearly indicated. (www.puc.edu)
http://www.puc.edu/Faculty/Gilbert_Muth/phot0039.jpg

Radicle: The embryonic root enclosed within a seed. (Raven, G-20) The radicle is attached to the hypocotyl inside the seed, and is the first organ to emerge from the germinating seed. (Campbell Reece, 7th Edition, pg 780)
Amanda M
Enhancement: The radical is made up of the root apical meristem and the root cap. It is located at the lower end of the hypocotyl. During seed germination, the radical is the first to emerge from the seed and it plants itself into the soil. (Biology of Plants, 7th edition, Peter H. Raven Pg. 507).
Ilustration of radical emerging from seed
Enhancement by Daniel Struthers(text and picture)
Root Nodules: Enlargements or swellings on the roots of legumes and certain other plants inhabited by symbiotic nitrogen-fixing bacteria. The root nodules of legumes consist of a relatively narrow cortex, which surrounds a large central zone containing both bacteroid-infected and uninfected cells. Vascular bundles, which radiate from the point of attachment of the nodule to the root, occur in the inner cortex. (
Biology of Plants, 7th edition, Peter H. Raven, pg. 657-658)

Nitrogen-fixing nodules on the roots of a legume.
http://www.bio.usyd.edu.au/DavidDay/root_nodules.jpgKevin Christie
Enhancement: Root nodules are usually host to the Rhizobium and Bradyrhizobium bacterias, which are extremely effective nitrogen fixers. There is a symbiotic relationship between these nitrogen fixing bacterias and the plants that they inhabit. The bacteria provides the plant with a form of nitrogen that it can use to make protein, while the plant in turn provides the bacteria with an energy source.Amanda M
Taproot system:Taproot system derives directly from the first root that emerges from a seed that is called the radicle. It forms a central root, the taproot, which grows vertically down into the soil. (http://www.botany.uwc.ac.za/sci_ed/grade10/anatomy/roots.htm) The taproot is larger in diameter with a limited amount of smaller branches extending laterally referred to as the lateral roots. Taproots generally grow deep into the soil to obtain nutrients and water. It functions for storage and transport, providing reserves of food in the form of a carbohydrate or for reaching water deep in the ground. (Taproot – Botany) Examples of taproot system plants are nut trees, carrots, radishes, parsnips and dandelions. (Master Gardener. Ohio State University Extension)
References: http://www.botany.uwc.ac.za/sci_ed/grade10/anatomy/roots.htm (Taproot – Botany) http://www.cactus-art.biz/note-book/Dictionary/Dictionary_T/dictionary_tap_root.htm
(Master Gardener. Ohio State University Extension) http://www.hcs.ohio-state.edu/mg/manual/botany.htm
-Nikki Heim
Enhancement
Most trees begin life with a taproot, but after one to a few years change to a wide-spreading fibrous root system with mainly horizontal surface roots and only a few vertical, deep anchoring roots. The taproot is larger in diameter than the lateral roots. Lateral roots, which originate from the pericycle, branch off from the taproot, and subsequent lateral roots can branch off other lateral roots. A typical mature tree 30-50 m tall has a root system that extends horizontally in all directions as far as the tree is tall or more, but well over 95% of the roots are in the top 50 cm depth of soil. http://www-plb.ucdavis.edu/labs/rost/Tomato/Roots/taproot.html
Mike Tomlinson
Enhancement- This picture shows the visual difference in a tap root (carrot) vs. the fibrous root which is an onion. www.botony.hawaii.edu Jess.E.W

Turgor Pressure: From the Latin word “turgor”, meaning “a swelling”; Turgor pressure refers to the hydrostatic pressure in a plant cell that results from osmosis/imbibition. As the water flows into the cell it swells and the water pushes against the cell wall. The pressure of the water against the cell wall is Turgor Pressure. The swelling of the cells due to increased Turgor pressure causes the embryo inside a seed to enlarge and eventually split the seed coat, allowing germination. (Biology of Plants, 7th Edition, Peter H. Raven, pg 75, G-25) (Biology, 6th Edition, Neil A. Campbell, pg 752, Glossary) –Tori Waites
Enhancement:
It is determined how much water is in the vacuole and resulting from osmotic pressure. Turgid plant cells contain more water than flaccid cells and exert a greater osmotic pressure on its cell walls. The pressure of each cell wall against surrounding cell results in stiffness that allows the plant to stay upright.

Ref:
http://www.ccr.buffalo.edu/etomica/app/modules/sites/Osmosis/Background1.html
Campbell, N. A., Reece, J. B. Biology: Sixth Edition; Benjamin Cummings: New York, NY, 2002; Vol. 1.
-Scott Horley
Compare & Contrast (first word alphabetical)
Adventitious roots/Feeder roots: Amanda Last
Adventitious root (syn. stem root)A root that arises from a stem, rather than from the primary root. http://glossary.gardenweb.com/glossary/adventitious_root.html Referring to a structure arising from an unusual place, such as buds at other places than leaf axils, or roots growing from stems or leaves. (Biology of Plants, Raven, G-1)
Some roots, called adventitious roots, arise from an organ other than the root—usually a stem, sometimes a leaf. They are especially numerous on underground stems. The formation of adventitious roots makes it possible to vegetatively propagate many plants from stem or leaf cuttings. (Britannica On-line encyclopedia).
Feeder Roots: Fine roots actively engaged in the uptake of water and minerals. The bulk of most feeder roots usually occurs in the uppermost meter of soil. In most trees feeder roots primarily occur in the upper 15cm of soil, which is normally richest in organic matter (Biology of Plants, Raven, p.529) The feeder, or fine, roots are similar to those of herbaceous vascular plants until, as they mature, they begin to undergo secondary growth(Britannica On-line Encyclopedia) Both the adventitious roots and feeder roots are part of the root system. Adventitious roots come do not come from the primary root but rather from a stem or bud. Feeder roots are finer and take part in absorbtion of water and minerals.
Enhancement(Adventitious roots/Feeder roots) Russ Fountain


(Above Left)This is an example of a monocot with adventitious roots.The roots are coming off stems in this instance, and are therefore called adventitious. (“Monocot or Dicot – Adventitious Roots” -
http://www.bridgewater.edu/~lhill/adventitiousroots.htm)
(Above Right)At the surface, feeder roots mix with lawn and shrub roots and compete for the water, oxygen and minerals that are more abundant near the surface. (“Tree Root Basics” Southern Nevada Water Authority -
http://www.snwa.com/html/land_trees_roots.html)
Adventitious roots/Fibrous root system (Dennis Murengi)
Adventitious roots: These are roots growing from stems or leaves. (Biology of Plants 7th Edition G-1)
Fibrous root system: A root system made up of many threadlike members of more or less equal length. (www.thefreedictionary.com)
http://www.thefreedictionary.com/fibrous+root Adventitious roots and fibrous roots are similar in the functions they perform. These functions include anchorage, storage, absorption, and conduction/transport. However, adventitious roots and fibrous root system differ in terms of how they from. Adventitious roots arise from the stems or leaves of plants. On the other hand the fibrous root system develops from adventitious roots arising form the plant’s stem. (www.thefreedictionary.com)
http://www.thefreedictionary.com/fibrous+root

The image on the left shows adventitious roots growing from a leaf, while the image on the right shows a fibrous root system.
Apical meristem/lateral meristem Apical meristem:
Tissue located at the tip of stems and roots where cells are continuously and repeatedly dividing. Cell division of apical meristems results in primary growth, increasing the length of stems and roots. (Raven, 7th ed.) (Biolog online: http://www.biology-online.org/3/7_meristems.htm)
Lateral meristem
Tissue located in the vascular cambium and cork cambium of stems and roots that are no longer elongating. Cell division in these tissues produces secondary growth, increasing the thickness of the plant body. (Raven, 7th ed.) (Biolog online: http://www.biology-online.org/3/7_meristems.htm)
Apical and lateral meristems are both tissues where cell division contributes to new growth in the plant body. Lateral meristems increase the diameter of stems and roots while apical meristems increase the length.
Katherine Loewen

This is an image of the apical meristem.
www.answers.com (Matti Harper)
This is an image comparing the apical and lateral meristems. www.agen.ufl.edu/.../lect/lect_15/22_2AT.GIF (Matti Harper)
Apical meristems/ Primary meristem The apical meristem is the growing tips of a root that is undifferentiated meristematic tissue. This is were new growth begins with new cells forming at the tips of the roots and shoots. From the primary meristem arise from the apical meristems that differentiate into three kinds of primary meristems including the protoderm, procambium, and ground meristem. Ref: http://arnica.csustan.edu/boty1050/Tissues/tissues.htm Scofield and Murray (2006). The evolving concept of the meristem. Plant Molecular Biology 60:v–vii -Scott Horley
Apical Meristem: Embryonic plant tissue in the tips of roots and the buds of shoots that supplies cells for the plant growth in length (Biology 7th Edition, Campbell and Reece, pg. 414) It is a mass of undifferentiated cells at the tip of the stem and is responsible for primary growth (http://users.rcn.com/jkimball.ma.ultranet/BiologyPages/F/Flowering.html).
Ground Meristem: The primary meristem, or meristematic tissue that gives rise to the ground tissues (Biology of Plants, Raven, pg. 498). It develops into the pith and the cortex and surrounds the procambium, which will extend into other regions of the embryo as embryogenesis continues (http://dictionary.reference.com/browse/ground%20meristem).
Summary: The apical meristem and ground meristem are similar in that they are both responsible for primary growth and are both parts of the meristematic regions of a plant. They differ in that apical meristems are part of the vascular tissue system and the ground meristem is part of the ground tissue system. They are also different in that the apical meristem is responsible for plant growth in length where the ground meristem develops into the pith and cortex. (Lacey Loewen)
Casparian Strip and Exodermis-Danielle Dornik
| Casparian Srip | Exodermis |
• Band of Primary cell walls • Contain suberin(fatty substance) • Stop water and minerals going though side walls • Is in Endodermal cells and Exodermal cells(Exodermal only in angiosperms) • Functions mainly in root hair zone-where water is taken up | • Outer layer of cortex • Band of Casparian strip (Agiosperms only!!) • Have radial and transverse walls • Located below epidermis • Reduces water loss for the root • Provides defence against microorganisms |
As shown above one can link the casparian strip functions to that of the Exodermis. It is important to remember that water can not go through the side walls. Also that the exodermis is only in agiosperms!
Biology of Plants, 7th edition, Peter H. Raven, Glossary pg. G-9
http://www.eeob.iastate.edu/classes/bio454/docs/404root104.pdf
Endodermis/Exodermis The exodermis is the outermost layer of the cortex, whereas the endodermis is the innermost layer of the cortex. Both cells are characterized by Casparian strip within radial and transverse wall. They both direct water through the protoplasts, where the exodermis reduces water loss and the endodermis filters the water reducing the risk if disease entering the stele. The endodermis is a single layer of cells and the exodermis is one or more cells in depth. (Biology of Plants 6th Edition)-Derek Burdikin
Sexual reproduction/ Asexual reproduction in plantsThe vast majority of plants reproduce sexually. This can be only means of reproduction or it can be in combination with one or more means of asexual reproduction. Sexual reproduction requires haploid spores from a second parent for fertilization.Asexual reproduction involves only one parent that produces genetically identical offspring. It is also known as vegetation reproduction because the clones are usually fragments of the parent plant. Examples of asexual reproduction are fragmentation and layering.Each method offers advantages in certain situations. Asexual means is best suited for stable, favoring environments where the clones will be as productive as the parent. If the conditions are changing or the parent plant is not best suited for the environment than sexual reproduction is beneficial. Kirsten (Biology 7th ed., Campbell & Reece, p. 781), (Biology of Plants, Raven, p 158)
Tap Root System/Fibrous Root System (Kevin Christie) Tap Root System: The primary root of a plant formed in direct continuation with the root tip or radicle of the embryo: forms a stout, tapering main root from which arise smaller, lateral roots. The older lateral roots are found nearer the base of the root (where the root and stem meet), and the younger ones nearer the root tip. Taproot systems generally penetrate deeper into the soil than fibrous root systems.( (Biology of Plants, 7th edition, Peter H. Raven, pg. 528 – 530) Fibrous root system: Root system in plants which no one root is more prominent than the others. Typically shallower than tap root system and the tenacity with which they cling to soil particle make such plants especially well suited as ground cover for the prevention of soil erosion. (Biology of Plants, 7th edition, Peter H. Raven, pg. 528 – 529)Transverse Wall/Radial Wall (Caitlin Deas) Both a
transverse wall and a
radial wall are anticlinal walls in the root endodermis. They are perpendicular to the root (Botany,
http://biology.jbpub.com/Botany/interactive_glossary_showterm.cfm?term=anticlinal%20wall). The transverse wall is at a right angle to the long axis of the root along the top of the cell, while the radial wall is at a right angle to the root’s long axis but on the side of the cell.
(University of Minnesota Extension)
http://www.extension.umn.edu/distribution/naturalresources/components/6413ch1.htmlLignin and suberin impregnate the transverse and radial walls to form the Casparian strip, making these walls important for blocking water from entering the cells without being filtered.
(The Ecotree)
http://www.bcb.uwc.ac.za/ecotree/root/rootA.htmBelow is an image of the transverse and radial walls showing the Casparian strip. The long axis of the root would be in the front of the picture, blocking the view of the cells.

(The Ecotree)
http://www.bcb.uwc.ac.za/ecotree/root/images/Endodermis.GIFVascular Cambium / Stele: Vascular Cambium – cylindrical sheath of meristematic tissue, the division of which produces secondary xylem and phloem (Raven, Biology of Plants, pg G-25). Stele – central cylinder, inside the cortex of the roots and stems in vascular plants (Raven, Biology of Plants, pg G-22), the vascular tissue inside the stem or roots (Campbell & Reece, Biology 7th ed. Glossary). Primary vascular tissues (primary xylem and phloem), and in some vascular plants the pith, make up the central cylinder, or stele, of the stem and/or root in the primary plant body (Raven, Biology of Plants, pg 371). Secondary growth of the xylem and phloem into the secondary xylem and phloem is caused by the activity of the lateral meristem called the vascular cambium which adds layers of vascular tissue (secondary xylem and phloem) (Campbell & Reece, Biology 7th ed. Pg 720). Stephanie Lauer Enhancement: (AarenS) Vascular cambium can be seen in picture below, note the location between the sapwood and secondary phloem.
(www.britannica.com)Stele shown below, comprised of phloem, cambium, xylem and pericycle.
www.apsnet.org Vascular Cylinder/Pericycle (Daniel Struthers): Both part of the vascular tissue system and originate from the procambium and found in the root system. They are both important parts of the stele. The Vascular Cylinder of the root system consists of primary vascular tissues and one or more layers of nonvascular cells. Consisting of the xylem and phloem, it is responsible for water and food transportation. The Pericycle completely surrounds the vascular tissues within the vascular cylinder. In most seed plants, lateral roots arise in the pericycle and also contributes to secondary growth in the vascular cambium.(Biology of Plants, 7th Edition, Peter H. Raven, Pg. 538) Illustration of Vascular cylinder/pericycle (www-plb.ucdavis.edu/.../Roots/rootorg.html) 